For cable-driven speedometers: the driven gear tooth count that makes your speedometer read true after a gear or tire change, and how far off the nearest available gear will be.
A cable-driven speedometer expects its cable to turn a set number of times per mile: 1,001 for most GM units and 1,000 for Ford. The driveshaft turns tire revolutions × axle ratio times per mile, and the drive and driven gears step that down to what the speedometer expects.
Gears only come in whole teeth, so the nearest gear is rarely perfect. The result shows how far off it will read so you can choose between the two closest gears.